Ray Romano

Manny (voice)

John Leguizamo

Sid (voice)

Denis Leary

Diego (voice)

Queen Latifah

Ellie (voice)

Simon Pegg

Buck (voice)

Seann William Scott

Crash (voice)

Josh Peck

Eddie (voice)

Chris Wedge

Scrat (voice)

Karen Disher

Scratte (voice)

Kristen Wiig

Pudgy Beaver Mom (voice)

Eunice Cho

Madison (Diatryma Girl) (voice)

Maile Flanagan

Aardvark Mom (voice)

Bill Hader

Gazelle (voice)

Joey King

Beaver Girl (voice)

Jane Lynch

Diatryma Mom (voice)

Clea Lewis

Start Mom (voice)

Carlos Saldanha

Dinosaur Babies / Flightless Bird (voice)

Claudia Besso

Additional Voice (voice)

Bea Miller

Additional Voice / ADR Group (voice)

Matt Adler

Additional Voice (voice)

Steve Alterman

Additional Voice (voice)

Holly Dorff

Additional Voice (voice)

David H. Kramer

Additional Voice (voice)

Selenis Leyva

Additional Voice (voice)

Devika Parikh

Additional Voice (voice)

Johnathan Tchaikovsky

Additional Voice (voice)

June Christopher

Additional Voice (voice)

Frank Welker

Momma Dino (voice) (uncredited)